Glossary

Aperture – a hole or thinning in structure of the pollen grain – distinct from ornamentation in form and function.

Baculate – ornamentation that is rod-like, base not constricted. Boat – pollen grain whose shape is boat- or kidney-shaped.

Clavate – sculpturing greater than 1 μm with constricted bases.

Colpus – an elongated aperture or furrow in the exine. Diporate – having two pores.

Disulcate – two elongated apertures at either pole of the grain.

Echinate – sculpturing with spine- or spike-like elements, between 1 and 3 μm in length.

Fossulate – tectum with furrow-like depressions.

Gemmate – sculpturing with elements constricted at the base, less than 1 μm.

Inaperturate – without pores, letes, sulci, or colpi. No apertures.

Irregular – pollen grain whose shape is not regular. Leptoma – a thinning in the exine of
some Pinaceae pollen grains.

Microechinate – sculpturing with spike-like elements, less than 1 μm in length.

Microreticulate – net-like ornamentation with the openings less than 1 μm in size.

Monad – pollen grains typically dispersed singly.

Monocolpate – an aperture type with a single colpus.

Monoporate – having one pore.

Monosulcate – a single elongated aperture located at the pole of the grain.

Oblate – pollen grain that is wider than it is tall.

Ornamentation – patterned surficial modification to the exine of a pollen grain.

Panto- /periporate – an aperture type where the pores are evenly distributed over the surface.

Perforate – tectum with holes or depressions.

Panto - /pericolpate – an aperture type with more than three colpi distributed over the surface.

Polyad – pollen grains typically dispersed in groups of greater than four.

Polyplicate – a pollen type with more than three deep folds and ridges.

Pore – a circular aperture or hole in the exine.

Prolate – pollen grain that is taller than it is wide.

Psilate – tectum or surface of the pollen grain that is smooth.

Rectangular – pollen grain that has a rectangular outline.

Reticulate – net-like ornamentation with the openings greater than 1 μm in size.

Rhombic/polyhedral – pollen grain that has several flattened faces.

Rugulate – ornamentation that has forms irregular ridges.

Scabrate – having raised, round surface elements less than 1 μm in size.

Spheroidal – pollen grain that is roughly spherical; round in outline.

Stephanocolporate – an aperture type with more than three colpi arranged longitudinally.

Stephanoporate – an aperture type with more than three pores distributed around the equator of the grain.

Striate – ornamentation that forms roughly parallel ridges.

Tetrad – pollen grains typically dispersed in groups of four.

Triangular – a pollen grain whose outline is triangular. Tricolpate – an aperture type with three colpi.

Tricolporate – with three pores, usually associated with three colpi.

Trilete – a three-pronged scar on a spore.

Triporate – with three pores.

Verrucate –surface sculpturing with raised round elements greater than 1 μm in size.

Vesiculate – a pollen type with bladders; common in Pinaceae.

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